Main Research Interest
Ecology, taxonomy and ultrastructure of nematodes

Background
A graduate in Zoology at Liverpool University, UK, a doctorate at the Liverpool School of Tropical Medicine on parasitological research in West Africa. A post doctorate Rockefeller Foundation Fellowship at Berkely, University of California. Visiting Professor at the University of Illinois. Lectured in Zoology at Liverpool University and since 1960 in Zoology at ANU until retirement in 1991.

Most Recent Publications
Nicholas, W.L. 2007. A new species of Trileptium (Nematoda, Thoracostomopsidae) from a sandy beach in southeastern Australia, with a key to species and observations on geographical distribution. New Zealand Journal of Marine and Freshwater Research 41: 335-344.

Nicholas, W. 2006. The meiofauna of a New South Wales sandy beach: an introduction to the meiofauna of Australian ocean beaches. Wetlands 23(2): 14-31.

Nicholas, W.L. 2002. Ultrastructure of the Head of Okranema eileenae Greenslade and Nicholas, 1991 (Thoracostomopsidae: Nematoda). Journal of Nematology 34(1) 50-61.

Nicholas, W.L. 2001. Seasonal variations in nematode assemblages on an Australian temperate ocean beach; the effect of heavy seas and unusually high tides. Hydrobiologia 464: 17-26.